KI population's brood were present in nests primarily from late spring to late summer.
WW populations brood occurred in mid-summer.
Egg laying began later in KI than in WW as this is
reflected by smaller ovary sizes during early spring
and fewer #'s of brood
For KI, few brood were found in the autumn and one
pupa was found in winter (possibly due to late
summer production when development times are
In KI + WW Multi-female colonies tended to occur
during the beginning of brood-rearing, but by the
end, the colonies were solitary female.